The Enzymatic Sentinel:
Alanine Aminotransferase
A structural and biochemical analysis of ALT. From cytosolic gradients to prognostic modeling in metabolic liver disease.
The 3,000-Fold Differential
Alanine Aminotransferase (ALT), EC 2.6.1.2, is defined not by its presence in serum, but by its overwhelming concentration within the hepatocyte. The biochemical gradient is stark: concentrations in the cytosol are approximately 3,000 times greater than in the extracellular space.
The P5P Dependency
ALT activity is catalytically dependent on Pyridoxal-5'-Phosphate (P5P), the active form of Vitamin B6. The enzyme exists as an inactive apoenzyme until it binds P5P to become the active holoenzyme.
In states of malnutrition (e.g., chronic alcohol use), P5P depletion leads to a "silent" liver injury where ALT levels remain artificially low despite active necrosis.
The Diagnostic Triad
The De Ritis Ratio
The ratio of AST to ALT serves as a temporal and etiological marker. The interpretation relies on the kinetics of clearance and the subcellular location of the enzymes.
- Ratio < 1 (ALT Dominant): Classic acute viral hepatitis and MASLD. Cytosolic release prevails.
- Ratio > 2 (AST Dominant): The alcohol signature. Driven by mitochondrial damage plus B6-induced ALT suppression.
- The Inversion (>1 in chronic disease): A marker of advanced fibrosis/cirrhosis. As hepatic blood flow changes, sinusoidal clearance of AST drops, causing it to accumulate.
Mechanism of Injury
Metabolic Dysfunction (MASLD)
The primary driver is Lipotoxicity. Free fatty acid accumulation overwhelms mitochondrial beta-oxidation, triggering oxidative stress. Treatment focuses on 7-10% weight loss to reverse histology.
Acetaminophen (APAP)
Saturation of sulfation pathways shunts metabolism to CYP2E1, producing toxic NAPQI. This depletes glutathione, causing rapid centrilobular necrosis.
In end-stage cirrhosis, ALT may normalize simply because the viable hepatocyte mass is depleted. Always cross-reference with synthetic function markers (INR, Albumin).
Clinical Takeaway
- New Normals: Accept lower cutoffs (M:33, F:25) to detect early metabolic disease.
- Low ALT: Values < 15 U/L in the elderly are a validated biomarker for sarcopenia and frailty.
- The Ratio: An inverted AST/ALT ratio in a Hepatitis C patient is a strong specific indicator of underlying cirrhosis.